New tribes (Paraphytini, Coptorhinini) proposed for five African or Afro-Oriental dung beetle genera formerly classified in the tribe Ateuchini Perty, 1830 (Coleoptera: Scarabaeidae: Scarabaeinae).

Owing to discrepancies between phylogeny and tribal phenetic classification in the subfamily Scarabaeinae, a number of new or old tribes have recently been described or revalidated to resolve the differences. The present paper continues this process…

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